http://www.sciencedaily.com/releases/2012/06/120613102132.htm
ScienceDaily (June 13, 2012) — Dogs can sometimes suffer sudden episodes of aggression without their owners understanding why. But, in many cases, the cause of these attacks can be pain that has never been diagnosed or treated. For the first time the study describes the characteristics of this irritability, which can make dogs violent and increase aggression in already conflictive individuals.
There are many factors that explain aggression in dogs: the conditions of the mother during gestation, the handling of the puppy in the neonatal phase, the age at weaning, the experiences of the animal during the socialising phase, diet, exercise, genetics and learning techniques based on active punishment during adulthood. However, aggressive behaviour also arises from the presence of pathologies and pain in the dog.
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